PI3Kalpha Translocation Mediates Nuclear PtdIns(3,4,5) P3 Effector Signaling in Colorectal Cancer
Details
Publication Year 2023-04,Volume 22,Issue #4,Page 100529
Journal Title
Molecular & Cellular Proteomics
Publication Type
Research article
Abstract
The canonical view of PI3Kalpha signaling describes phosphatidylinositol-3,4,5-trisphosphate (PtdIns(3,4,5)P(3)) generation and activation of downstream effectors at the plasma membrane or at microtubule-bound endosomes. Here, we show that colorectal cancer (CRC) cell lines exhibit a diverse plasma membrane-nuclear distribution of PI3Kalpha, controlling corresponding levels of subcellular PtdIns(3,4,5)P(3) pools. PI3Kalpha nuclear translocation was mediated by the importin beta-dependent nuclear import pathway. By PtdIns(3,4,5)P(3) affinity capture mass spectrometry done in the presence of SDS on CRC cell lines with PI3Kalpha nuclear localization, we identified 867 potential nuclear PtdIns(3,4,5)P(3) effector proteins. Nuclear PtdIns(3,4,5)P(3) interactome proteins were characterized by noncanonical PtdIns(3,4,5)P(3)-binding domains and showed overrepresentation for nuclear membrane, nucleolus, and nuclear speckles. The nuclear PtdIns(3,4,5)P(3) interactome was enriched for proteins related to RNA metabolism, with splicing reporter assays and SC-35 foci staining suggesting a role of epidermal growth factor-stimulated nuclear PI3Kalpha signaling in modulating pre-mRNA splicing. In patient tumors, nuclear p110alpha staining was associated with lower T stage and mucinous histology. These results indicate that PI3Kalpha translocation mediates nuclear PtdIns(3,4,5)P(3) effector signaling in human CRC, modulating signaling responses.
Publisher
Elsevier
Keywords
Humans; *Phosphatidylinositols/metabolism; Phosphatidylinositol Phosphates/metabolism; Signal Transduction; Cell Nucleus/metabolism; *Colorectal Neoplasms/metabolism; PI3Kalpha; PtdIns(3,4,5)P(3) interactome; colorectal cancer; nuclear import pathway; pre-mRNA splicing
Department(s)
Medical Oncology
PubMed ID
36931626
Terms of Use/Rights Notice
Refer to copyright notice on published article.


Creation Date: 2023-07-11 06:23:30
Last Modified: 2024-07-16 04:37:01

© 2024 The Walter and Eliza Hall Institute of Medical Research. Access to this website is subject to our Privacy Policy and Terms of Use

An error has occurred. This application may no longer respond until reloaded. Reload 🗙